Caldasia

DESCRIPTION

Measurements. See table 1.

Dorsum. DS outline epsilon, type 2. Ocularium elliptical and medium, without median depression, tuberculated, located medially on the carapace. Mesotergum (abdominal scutum) delimited, divided into four areas, each one with a pair of paramedial large granules (Figs 1b, d, e, 2a-b). DS widest at level of area II. Area I divided into two halves with some granules; areas II--IV undivided and with some granules (Figs 1b, 2a-b). Posterior border of scutum curved (Figs 1a-b). Free tergites I--III with a row of granules (Figs 1b, d, 2a-b).

Venter. Stigmatic area with a few granules and a posterior row of conspicuous granules. Stigmata oval and concave (Fig. 1c). Coxa I with a curved row of tubercles increasing in size distally, and one proximal, large, and trapezoidal tubercle on the anterior margin of the coxa I, with a medium-sized tubercle at its base, and one small tubercle at the proximal face (Fig. 2c); coxa II longer than coxa I; coxa III longer than I and II; coxa IV backward (Fig. 1c).

Chelicera. Basichelicerite (segment I) trapezoid with well-marked bulla (Figs 1b, d); three large tubercles on ectal face, two large tubercles on posterior margin, and two medium-sized tubercles on the dorsomedial face (Figs 1b, d, 2a). Chelicera swollen (Figs 1, 2a, d). Anterior region of the hand with setiferous tubercles of different sizes that go from the medial region of the hand to the base of movable and fixed fingers. Fixed finger with the inner surface finely grooved. Movable finger with one trapezoid sub-basal tooth and the inner surface of the distal portion dentated (Fig. 2d).

Pedipalps. Trochanter with one ventral subapical tubercle. Femur with a ventroectal row of four setiferous tubercles, the two basalmost larger than the two distalmost, and one medium ventromesal setiferous tubercle in the apical portion (Figs 1c, 2e). Patella with one large mesal setiferous tubercle. Tibia ectal iIIi, mesal IIi. Tarsus ectal IIi, mesal IIi (Fig. 2e).

Legs. Leg I filiform, legs I--IV straight and smooth. Leg IV thicker than the others; coxa IV with a large dorso-distal tubercle dorsally projected (Figs 1a, d-e, 2b); Fe IV length almost three and a half times DS length (Fig. 1a). Tarsus IV without tarsal process, claws smooth. Tarsal counts: 7(3)-8(3)/15(3)-?/7-7/8-? (holotype), 7(3)-7(3)/15(3)-?/7-7/6-6 (paratype).

Penis. LP small (width twice the height) and apically depressed with anterolateral blunt and not dorsoapically pointed swollen tips (Figs 2f, g). Malleus with two pairs of branched MS-A located in the dorsal half, MS-A2 located far from the base of the stylus; one pair of branched MS-B on the ventral side; MS-C absent; two pairs of medium-sized MS-D1--D2 located close to each other and near the stylus on a keel between the dorsal region of the LP and the base of the stylus; MS-E2 large and branched, MS-E1 medium-sized, conical, and located slightly distal to MS-E2 on the ventral side of the LP (Figs 2f, g). Stylus elongated, mostly straight, and surpassing the LP; connective tissue present between the sharp DP and the stylus, slightly corrugated, with irregular peaks on the apical medial region (Figs 2f, g).

Color (in alcohol). Background of DS, chelicerae, and mesotergal paired granules Moderate Yellow (87). Lateral borders of DS, free tergites, and mesotergal areas around the paired granules Dark Grayish Brown (62). Pedipalpal segments of different colors: trochanter, the first half of the femur, dorsal spot of the tibia, tarsus, and claw Dark Grayish Brown (62); distal half of femur, patella, and tibia Pale Yellow (89). Legs I--IV Dark Grayish Brown (62) (Fig. 1).

Female. Unknown.

Distribution and habitat. The species is only known from the forests of the lowlands of Meta department (Acacías municipality) (Fig. 5), in the limits between the Cordillera Oriental montane forests and the ecoregion of the Apure-Villavicencio dry forests (Olson et al. 2001).

Etymology. The species is named after the late entomologist German Amat García, for his great teaching contribution to Colombian entomology. Amat is the Catalan version of the Latin surname Amatus, meaning the loved one. Dilectus, in Latin, has the same meaning.

DESCRIPTION

Measurements. See table 1.

Tabla 1: Measurements (in mm) of body and appendages of Vima dilectus sp. nov. and Vima bellator sp. nov.

Figura 1: Photographs of the habitus of Vima dilectus sp. nov. (ICN-Ao-411.1), male holotype: a. panoramic, b. dorsal, c. ventral, d. lateral, e. frontal. Scale bars: a = 3 mm, b, c, e = 1 mm, d = 0.5 mm.

Diagnosis. Segments of pedipalp concolor (as in V. insignis; in V. panita and V. dilectus the distal femur, patella, and tibia are whitish). Anteroproximal tubercle of coxa I with three marked peaks (smooth in the rest of the species). Low ocularium (high in the rest of the species). Peaks on the connective tissue between DP and stylus higher at medial region (same height in V. bellator; mostly smooth and with just one medial peak in V. panita; tissue absent in V. insignis).

Dorsum. DS outline epsilon, type 2. Ocularium medium, located medially on the carapace, without median depression and smooth. Mesotergum delimited and divided into four areas each one with a pair of paramedial low granules (Figs 3b, d, e, 4a-b). DS widest at level of area II. Area I divided into two halves, ornate with some granules; areas II--IV undivided and with some granules (Figs 3b, 4a-b). Posterior border of scutum convex (Figs 3b, 4a). Free tergites I--III with a row of granules (Figs 3b, d, 4a-b).

Venter. Stigmatic area with few granules. Stigmata oval and transverse (Fig. 3c). Coxa I with a sigmoidal row of tubercles increasing in size distally, one proximal, large, tricapitated tubercle on the anterior margin, and one medium-sized tubercle on the posterodistal region (Fig. 4c); coxa II longer than coxa I; coxa III slightly longer than I and II; coxa IV backward (Fig. 3c).

Chelicera. Basichelicerite trapezoid, with well-marked bulla (Figs 3b, d); two large tubercles on the ectal face, three large tubercles on the posterior margin, and some medium-sized tubercles on the dorsomedial face (Figs 3b, d, 4a). Chelicera swollen (Figs 3, 4a). Anterior region of the hand with small setiferous tubercles on the medial region, and a group of medium-sized/large tubercles close to the base of the movable and fixed fingers. Fixed finger with the inner surface finely grooved. Movable finger with one trapezoid sub-basal tooth, and with the inner surface on the distal portion dentated (Fig. 4d).

Figura 2: Drawings of Vima dilectus sp. nov. Male holotype (ICN-Ao-411.1): a. Habitus in dorsal view, b. same, lateral view; c. coxa I, ventral view; d. chelicera, frontal view; e. left pedipalp, mesal view. Genitalia of male paratype (ICN-Ao-411): f. lateral view, g. dorsal view (the letters A-D refers to the chaetotaxy system of Kury and Villarreal (2015)). Scale bars: a = 1 mm, b-e = 0.5 mm, f, g = 0.1 mm.

Figura 3: Photographs of the habitus of Vima bellator sp. nov. (ICN-Ao-575), male holotype: a. panoramic, b. dorsal, c. ventral, d. lateral, e. frontal. Scale bars: a = 3 mm, b, d = 1 mm, c, e = 0.5 mm.

Pedipalps. Trochanter with one ventral, subapical, and medium-sized tubercle. Femur with a ventroectal row of four setiferous tubercles, the two basalmost larger than the two distalmost, and one medium-sized ventromesal setiferous tubercle on the distal portion (Figs 3c, d, 4e). Patella with one large, mesal, setiferous tubercle. Tibia ectal iIIi, mesal IIi. Tarsus ectal IIi, mesal IIi (Fig. 4e).

Legs. Leg I filiform, legs I--IV straight and smooth. Leg IV thicker than the others; coxa IV with one small dorso-distal tubercle dorsally projected (Fig. 3a); Fe IV length almost four times DS length (Fig. 3a). Tarsus IV without tarsal process, claws smooth. Tarsal formula of holotype: 7-7/?-15/7-7/7-7.

Penis. LP small (width twice the height) and apically depressed, with an anterolateral blunt and not swollen tips dorsoapically pointed, and with a wide distal cleft (Figs 4f, g). Malleus with two pairs of branched MS-A located almost dorsally. MS-A2 located far from the base of the stylus; one pair of branched MS-B; MS-C absent; two pairs of medium-sized MS-D close to each other and near to the stylus. MS-D1--D2 located on a keel between the dorsal region of the LP and the base of the stylus; MS-E2 large and branched, MS-E1 medium-sized, conical, and located slightly distal to MS-E2 on the ventral side of the LP (Figs 4f, g). Stylus elongated, mostly straight, and surpassing the LP; connective tissue present between the sharp dorsal process and the stylus, slightly corrugated, with irregular peaks on the apical medial region (Figs 4f, g).

Figura 4: Drawings of Vima bellator sp. nov. Male holotype (ICN-Ao-575): a. Habitus in dorsal view, b. same, lateral view; c. coxa I, ventral view; d. chelicera, frontal view; e. right pedipalp, mesal view. Genitalia of male paratype (MPUJ_ENT 0064488): f. lateral view, g. ventral view (the letters A-D refers to the chaetotaxy system of Kury and Villarreal (2015)). Scale bars: a = 1 mm, b-e = 0.5 mm, f, g = 0.1 mm.

Color (in alcohol). Background of DS and chelicerae Moderate Yellow (87). Lateral borders of DS and free tergites Dark Grayish Brown (62). Mesotergal areas Moderate Brown (58) with transverse bands Moderate Yellow (87), pedipalps, and legs I-IV Moderate Brown (58) (Fig. 3).

Sexual dimorphism: Female with not hypertelic chelicera.

Distribution and habitat. The species is known from the forests of the lowlands of Santander department (Cimitarra and Puerto Parra municipalities) (Fig. 5) situated in the ecoregion of the Magdalena Valley montane forests (Olson et al. 2001).

Etymology. This species is named after the Colombian entomologist German Amat García. The name German, the Slavic form of Herman, is a masculine given name from an ancient Germanic name formed by harja (army) and mann (man), meaning warrior. Bellator, in Latin, has the same meaning.